Question

请问以下句子中的主句谓语是不是被省略了？谓语在哪里？

However, a question that remains as to whether the functions of MKP1 and MPK6 are only required during initial PAMP responses, or whether they function at multiple points in the response pathway.

Answer 1

这是并列复合句，主谓齐全，没有省略。其结构如下：
第一分句：a question are only required during initial PAMP responses
〔主语〕a question that remains as to whether the functions of MKP1 and MPK6 (that 引导的是定语从句)
〔谓语动词〕are required
〔程度状语〕only
〔时间状语〕during initial PAMP responses
第二分句：or whether they function at multiple points in the response pathway.
〔主语〕they
〔谓语动词〕function
〔地点状语〕at multiple points in the response pathway

Answer 2

所谓从句，它得是个句子，所以都有自己的主语和谓语。基本结构是: 主语从句（其中包含从句自己的主语+从句谓语） + 主句谓语比如： [What I just said] is really important. [That we are not prepared for the future] concerns us.